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It remains unknown whether a sucrose transporter mediates sugar signaling.Here we report that the Arabidopsis (Arabidopsis thaliana) sucrose transporter SUT4, but not SUT1 nor SUT2, interacts with five members of the Arabidopsis cytochrome b5 (Cyb5) family, and sucrose represses the interaction between SUT4 and a Cyb5 member Cyb5-2/A.And more data provide evidence that the sucrose transporter SUT4 interacts with Cyb5 to positively mediate sucrose and glucose signaling in the sucrose/glucose-induced delay of seed germination.Furthermore, we needs detailed studies to conform how do SUT4 and Cyb5-2 function in the sucrose-and glucose-induced delay of seed germination in the future.