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The beetles(Coleoptera)are known for the ability to effectively fold their hind wings beneath the stiff protective elytra and unfold them immediately.These foldable hind wings have several significant properties that can be explained by analogizing its structural designs to origami models.This study reveals the relationship between folding-unfolding behavior and the mountain-valley fold creases distributing on the wings.First,we examine the following origami properties: flat foldability,folding directions and vertex concavity of single vertex crease pattern.We can then compare these structures with some commonly used origami model,producing standard samples,and applying stretching test to determine their fold-unfold trait.The orientation of micro-pleats distributed on the wing which can be described by the order parameter which is related to mechanical properties of different parts of the wings.These features can be used to improve the design of bio-inspired materials and thin films which possess space efficiency and fatigue resistance.