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The interaction between plants and plant-growth-promoting rhizobacteria (PGPR) is a complex,reciprocal process.On one hand, plant compounds such as carbohydrates and amino acids serve as energy sources for PGPR [1].On the other hand, after colonization, PGPR promote plant growth by synthesizing plant hormones and increasing mineral availability in the soil, and control plants diseases by producing antibiotics and inducing the resistance of plants [2-4].