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TCP(TB1,CYC and PCFs)genes encode putative transcription factors,which contain a basic helix-loop-helix domain referred to as the TCP domain and play an important role in plant development.During zygomorphic(bilaterally symmetrical)floral development,it has been shown that the CYC-like TCP genes are key regulators recruited independently in several species to control dorso-ventral(DV)identities.In papillionoideae,the model legume Medicago truncatula,provides a good system to investigate the mechanism in the control of floral asymmetry or the zygomorphic development.