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Myristicaceae is distinctive in having dioecious, unisexual flowers with stamens fused into a synandrium, which consists of a single whorl of sessile anthers borne around a sterile central column.The peculiar androecial features with the greatest variation have played an essential role in the definition of genera and represent key characters for their identification.To depict the androecium diversity and better understanding the origin and evolution of synandria in Myristicaceae, floral ontogeny and anatomy of staminate flowers of several development phases in Horsfieldia and Myristica were investigated respectively, using scanning electron microscopy and light microscopy.