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Xanthomonas oryzae pv.oryzicola, the causative agent of bacterial leaf streak,injects a plethora of effectors through the type Ⅲ secretion system (T3SS) into rice cells to cause disease.The T3SS, encoded by the hrp-hrc-hpa genes, is essential for the pathogen to elicit the hypersensitive response (HR) in nonhost tobacco and for pathogenicity in host rice.