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Cotton is the leading fiber crop in the world,producing the most prevalent natural fibers used in textile industry.Cotton fibers are single cells derived from outer epidermis of ovules.Their growth and development consist of four overlapping stages:fiber initiation,cell elongation,secondary cell wall deposition,and maturation.Fiber cells from G.hirsutum are 1000 to 3000 times longer than they are wide.With highly elongated structure composed almost of cellulose,cotton fibers are considered as an ideal model for studies of plant cell elongation and cell wall biogenesis.