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Nature consists primarily of living materials,many of which are pathogenic microorganisms capable of killing and converting other organisms into copies of themselves [1,2].For self-defense,eukaryotes have gradually evolved immune systems,including the innate and adaptive immune systems [3-7].Innate immunity has been found in plants,fungi,and metazoans;while jawed vertebrates have an evolved adaptive immune system in addition to the innate immunity defense [8].