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Dear Editor,Upon recognition of pathogen effectors,plant resistance (R) proteins trigger strong defense responses that restrict the growth and spread of pathogens.Most R proteins are nucleotide-binding (NB) and leucine-rich repeat (LRR) domain-containing proteins (NLRs) sharing structural similarity with animal NOD-like receptors (Dodds and Rathjen,2010).SNC1 (Suppressor of npr1,constitutive 1) is an Arabidopsis Toll/interleukin-1 Receptor (TIR)-type NLR that was originally identified through a gain-of-function autoimmune mutant,snc1,from a forward genetic screen.