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２．托拉综合症病毒（ＴＳＶ） 根据其病毒颗粒结构（无囊膜的二十面体，３０ｎｍ直径，带胞质复制体，一个单链ＲＮＡ基因组），ＴＳＶ被归类为细小核糖核酸病毒 （ｐｉｃｏｒｖｉｒｕｓ）。 ＴＳＶ于１９９２年首次发现于瓜亚基尔（Ｇｕａｙｏｇｕｉｌ）湾托拉河口附近的对虾
2. Toray Syndrome Virus (TSV) TSVs are classified as picorviruses based on their viral particle structure (non-enveloped icosahedron, 30 nm diameter, with cytoplasmic replicon, a single stranded RNA genome) . The first discovery by TSV of shrimp near the Tora Estuary in Guayoguil Bay in 1992