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Two new genera and species of fossil Palaeontinidae are described from Daohugou Village, Inner Mongolia.China:Cladocossus undulatus gen.et sp.nov.and Cricocossus paradoxus gen.et sp. nov.Both new genera are described based on well-preserved forewings.This discovery confirms the high diversity of palaeontinids during the Middle Jurassic.Both specimens have interesting modal structures which are new to Palaeontinidae:M five-branched and M3 with two branches.Based on this character,wing structural characteristic is discussed.