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云南禄劝中泥盆世海口组为以碎屑岩为主的滨海相含煤沉积，地层中产有丰富的鱼类、植物和双壳类化石；在个体古生态学研究的基础上，根据动植物群的组合特征及分布规律，识别出２个化石群落组合．将化石生态群落的分布特征及埋葬规律同煤岩学、沉积环境分析相结合，全面分析了角质残植煤的形成过程．
The Haikou Formation of the Middle Devonian in Luquan, Yunnan Province is a coal-bearing marine facies sedimentary facies mainly composed of clastic rocks. The strata are rich in fish, plants and bivalve fossils. On the basis of individual paleontology research, Flora characteristics and distribution of the combination of two fossil communities identified. Combining the distribution characteristics of fossil communities and burial laws with coal petrology and sedimentary environment analysis, the forming process of cutanous coal was comprehensively analyzed.