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Polycomb group (PcG) proteins are gene repressors that help to maintain cellular identity during development via chromatin remodeling.Fertilization-independent endosperm (FIE), a member of the PcG complex, operates extensively in plant development, but its role in rice is not yet fully investigated.We report here the isolation and characterization of a PcG member in rice, which we designated OsFIE2 for Oriza sative Fertilization-Independent Endosperm 2.OsFIE2 is a single-copy gene in the rice genome and shows a universal expression pattern.