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Regular chromosome segregation into two daughter cells is possible with a properly ordered bipolar division spindle.In animals and lower eukaryotes,centrosome structures are spindle pole organizers.Before the beginning of division they double,nucleate star MTs,and then separate by means of interaction and mutual sliding of antiparallel MT bundles of opposite stars.However,in higher plant cells,the centrosome and spindle polar organizers are not identified as cellular morphological structures.The principles of plant spindle formation are still not completely understood.