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Emergence of pathogenicity within a large and diverse saprophytic genus,such as Streptomyces,is an important evolutionary event.Here we use comparative evolutionary analysis of streptomycete genomes to assess the gene content thai defines the genus (core-genome),and identify genes associated with a pathogenic lifestyle (patho-genome) The genomes of (11) streptomyeetes.including the three plant pathogens S.scabies 87-22,S.turgtdiscabies Car8 and S.ipomoeae 91-03.