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Shewanella oneidensis,a facultative γ-proteobacterium,is extensively studied for redox transformations.Now it is becoming a research model for many other studies such as biofilm formation,respiration diversity,fatty acid metabolism and motility.S.oneidensis is motile by a single polar flagellum.The flagellar genes are organized into a four-tiered transcriptional system.However,it is clear that the whole flagellar system is complex and there may be other regulator elements involved in flagellar transcription,assembly and function.