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Resistance to carbendazim and other benzimidazole fungicides in Gibberella zeae is conferred by point mutations in theβ2-tubulin gene, a gene that plays an important role in spindle assembly.The spindle assembly checkpoint is a cellular surveillance system that ensures the fidelity of chromosome segregation, which is critical for maintaining genomic stability.In this study, we isolated and characterized two predicted checkpoint protein-encoding genes (Mad2 and Bub1) in G.zeae and designated them as FgMad2 and FgBub1, respectively.