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Far-red insensitive 219 (FIN219), a member of the GH3 gene family in Arabidopsis, is involved in phytochrome A-mediated far-red (FR) light signaling and the same locus as JAR1.Previous genetic studies revealed that FIN219 acts as an extragenic suppressor of COP1.The molecular mechanism, however,underlying the suppression of COP1 activity remains unknown.