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Rice is a model organism for studying the mechanism of cell wall biosynthesis and remolding in Gramineae.Cellulose, consisting of unbranched β-1,4-glucans, is one of the most abundant polymers on earth and has the simplest structure.However, its biosynthesis and deposition are very complicated and controlled by many factors.Using forward genetic approach,our group has identified several genes participating in cellulose formation and deposition.Mutations of these gene cause deficient cellulose biosynthesis and abnormal mechanical strength.BC3 is dynamin related 2B protein, possibly involved in cellulose synthase catalytic subunits (CESAs) trafficking.BC1l, rice CESA4 protein, directly catalyzes cellulose synthesis in secondary cell wall and is critical for plant growth.bc6-2 is another missense mutant in CESA9.It forms the cellulose synthase complex with CESA4 and CESA7 to participate secondary wall cellulose biosynthesis in rice.Here, we provide new evidence for the biological functions of the three genes in cellulose biosynthesis.All those findings provide insights into the roles of cellulose in rice plants.