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Brassinosteroids (BRs) are known to have a key role in regulating plant growthand development.BRs are sensed at the cell surface by the leucine-rich repeat(LRR) domain of the membrane receptor kinase BRASSINOSTEROIDINSENSITIVE I (BRI1).However,it was not yet clear how steroid binding at thecell surface would activate the cytoplasmic kinase domain of the receptor.