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How genomic composition is shaped by isolation,gene flow and selection? We sought for answers by re-sequencing 33 genomes of Sonneratia alba,which is a widespread mangrove species in the Indo-West Pacific(IWP)region.The results of comparative genomic analyses indicated low genetic diversity within populations and strong sequence divergence among populations from four major oceanic regions,i.e.the South China Sea,the East Indian Ocean,northwestern Australia and eastern Australia.